Overview
- A peer-reviewed field study in Ecology Letters documents an extreme sex ratio on Golem Grad with up to an average of 19 males per adult female.
- Males repeatedly ram, bite, mount, and stab females with a sharp tail spur, leaving roughly three quarters of females with genital injuries.
- Harassed females retreat to cliff edges, with documented falls including a GPS-tagged adult later found dead below a cliff.
- Compared with a nearby mainland population, females on the island show poorer body condition, reproduce less often, and have lower annual survival.
- The origin of the skew remains unresolved, with the Macedonian Ecological Society noting the cause is unknown and a human introduction hypothesis unproven.